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Michael P. Weitzman, The Evolution of Manuscript Traditions, Royal Statistical Society. Journal. Series A: General, Volume 150, Issue 4, July 1987, Pages 287–304, https://doi.org/10.2307/2982040
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Summary
The originals of ancient literary works gave rise to copies. These manuscripts were often copied in turn; often, too, manuscripts eventually perished. The editor of an ancient work must consider the relations among the manuscripts, extant and lost, which transmitted the text, and the structure of such manuscript populations in general. For example, most family-trees reconstructed by editors for the manuscripts of ancient works show exactly two main branches; is this to be expected, or due to some flaw in methods of reconstruction? Such questions are approached by modelling the evolving manuscript population through a birth-and-death process, with illustrative data for Greek and Latin literature.