Model: how increasing meristem size can promote organ permutations. (A) Graphical definition of an organ permutation, based on silique positions on the stem (left panels): non-canonical divergence angles (red) are generated in the phyllotactic sequence (right panel). (B) Theoretical impact of a modulation of meristem size on the sequence of organ emergence in the SAM, assuming that organ inhibitory fields are not scaling to meristem size and that no change of phyllotactic mode occurs.