Identification of possible mechanisms underlying natural variation in leaf morphology. A, Correlations of 35 PHMF-based phenotypic values with six traditional multivariate leaf phenotype values in P. tomentosa. Color scale represents correlation coefficient. B, Manhattan plot displaying genome-wide association study (GWAS) results for the PHMF-based phenotypic values in 435 P. tomentosa and 505 P. simonii unrelated individuals. C, Genetic effect of PsiYAB11 (SNP16-4788525) on leaf traits and transcript abundance. The number of each genotype is given above the graph. Middle line indicates the median. Box shows the 5th to 95th percentiles. Whiskers indicate the interquartile range. Dots are outliers.