Distribution of DNA gyrase across the archaeal domain. The complete (>90%) and partial presence of gyrA and gyrB genes in a taxon are indicated with full black circle and half circle, respectively. The empty circle indicates the absence of DNA gyrase genes. It should be noted, however, that absence of genes in uncultured taxa may be due to genome incompleteness. Asterisks refer to lineages that were newly discovered or expanded (at class taxonomic level) since the last survey. The phylogenetic relationship between the four major groups is shown schematically, the uncertain position of DPANN in archaeal phylogeny is indicated with a dotted line.