This article featured a typographical error in the title when first published online (29 May). “differenciation” has been corrected to “differentiation”. The Publisher apologizes to the authors and to readers for this error, which was introduced during production of the article. In addition, in the ‘Methods’ section, the text “we observed pairs defending territories no larger than 100 m2” has been corrected to read “we observed pairs defending territories no larger than 100 x 100 m”. This error in the original manuscript does not affect the results of conclusions of the study.

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